On the Dynamics of Exploited Fish Populations
Among the fishes, a remarkably wide range of biological adaptations to diverse habitats has evolved. As well as living in the conventional habitats of lakes, ponds, rivers, rock pools and the open sea, fish have solved the problems of life in deserts, in
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Raymond Beverton and Sidney Holt at work in the Fisheries Research Laboratory, Lowestoft during 1949. Ray Beverton (left) can be seen working next to a 3-dimensiona! cardboard model of a yield isopleth diagram a novel concept at the time. Sidney Holt can be seen operating a hand-Brunsviga calculating machine, the 1949 equivalent of a computer but requiring more effort to use. 5
On the Dynamics of Exploited Fish Populations Raymond J . H . Beverton Emeritus Professor of Fisheries Ecology University of Wales and
Sidney J . Holt Senior Scientific Advisor to the International Fund for Animal Welfare
The investigations described in this book were undertaken during the years 1947-1953, during the first half of which both authors were on the staff of the M A F F Fisheries Research Laboratory, Lowestoft, Suffolk, U K . The MS was submitted for publication by HMSO in 1954.
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S P R I N G E R - S C I E N C E + B U S I N E S S M E D I A , B.V.
First edition 1957 Reprinted 1965 Facsimile reprint 1993 © 1957, 1993 Springer Science+Business Media Dordrecht Originally published by Chapman & Hall in 1993 Softcover reprint of the hardcover 1st edition 1993 Typeset in Imprint ISBN 978-94-010-4934-4
ISBN 978-94-011-2106-4 (eBook)
DOI 10.1007/978-94-011-2106-4 Apart trom any fair dealing tor tne purposes ot researcn or private study, or criticism or review, as permitted under the U K Copyright Designs and Patents Act, 1988, this publication may not be reproduced, stored, or transmitted, in any form or by any means, without the prior permission in writing of the publishers, or in the case of reprographic reproduction only in accordance with the terms of the licences issued by the Copyright Licensing Agency in the U K , or in accordance with the terms of licences issued by the appropriate Reproduction Rights Organization outside the U K . Enquiries concerning reproduction outside the terms stated here should be sent to the publishers at the London address printed on this page. T h e publisher makes no representation, express or implied, with regard to the accuracy of the information contained in this book and cannot accept any legal responsibility or liability for any errors or omissions that may be made. A catalogue record for this book is available from the British Library Library of Congress Cataloging-in-Publication data available Cover illustration. The graph shows the response of long-term yield of North Sea haddock (Melanogrammus aeglefinus L . ) to various levels of fishing mortality rate. Based on singlespecies models incorporating stock-and-recuitment, both alone and combined with densitydependent growth (from Beverton and Holt, 1957; Figs 18.6 and 18.17) X-axis = Fishing mortality coefficient (F) Y-axis = Average long-term yield Blue lines = stock and recruitment models Green lines = stock and recruitment models combined with density-dependent growth Dashed line = constant parameter model (for comparison) Dotted area = zone of F giving the highest average yield Hatched area = theoretical extinction zone, if these h
