Rare Nematoceran Dipterans (Insecta: Diptera) from the Khasurty Locality, Transbaikalia
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Nematoceran Dipterans (Insecta: Diptera) from the Khasurty Locality, Transbaikalia E. D. Lukashevich* Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow, 117647 Russia *e-mail: [email protected] Received December 28, 2019; revised January 16, 2020; accepted January 16, 2020
Abstract—New taxa of Nematocera are described from the Lower Cretaceous Khasurty locality (Western Transbaikalia, Russia). These are Praearchitipula ribeiroi sp. nov., P. kaluginae sp. nov., P. podenasi sp. nov. (Pediciidae), ?Protanyderus madrizi sp. nov. (Tanyderidae), Eoptychoptera fasbenderi sp. nov. (Ptychopteridae), Gilkania transbaikalica gen. et sp. nov. (Chironomidae), Amorimyia robusta gen. et sp. nov. (Anisopodidae). Keywords: Pediciidae, Tanyderidae, Ptychopteridae, Chironomidae, Anisopodidae, Mesozoic, Lower Cretaceous, new taxa, SEM DOI: 10.1134/S0031030120060040
INTRODUCTION In 2003, geologists from the Buryatgeocenter (Ulan-Ude) discovered the Khasurty Mesozoic insect locality in Western Transbaikalia, and insects collected from there were passed by O.R. Minina to the Paleontological Institute of the Russian Academy of Sciences (PIN). Over the past years, the Arthropod Laboratory of PIN has conducted several expeditions to this locality (in 2004, 2009, 2014 and 2019). The site produced abundant fossils, while Diptera turned out to be the dominant order of insects, about 2500 imprints are housed at the collections of PIN. Description of Diptera from this locality began recently. So far only representatives of a few rare families of Nematocera have been described, including Axymyiidae (Blagoderov and Lukashevich, 2013), Bolitophilidae (Greenwalt and Blagoderov, 2019), Kaluginamyiidae (Lukashevich et al., 2020), some Ptychopteridae (Lukashevich, 2019), and Perissommatidae (Lukashevich and Blagoderov, 2020). This article continues a series of systematic descriptions of dipterans from Khasurty, rarely found in the Mesozoic fossil record. All specimens described below come from the Khasurty locality, located on the left bank of the Khasurty River in its middle course in the Zakamensk region of Buryatia (50°21′ N; 103°38′ E); deposits belong to the lower part of the Lower Cretaceous Gusinoozerskaya Group. The type material is housed in the A.A. Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow.
This work uses the previously adopted venation nomenclature and Nematocera system (Shcherbakov et al., 1995; Lukashevich and Ribeiro, 2019). Scanning electron photography of the uncoated imprints were taken using a Tescan Vega XMU microscope equipped with reverse (BSE) and secondary electron (SE) detectors. SYSTEMATIC PALEONTOLOGY Infraorder Tipulomorpha Family Pediciidae Osten-Sacken, 1860 Subfamily Pediciinae Osten-Sacken, 1860
R e m a r k s. This subfamily includes the overwhelming majority of the living genera of this small family (11 out of 12). It is characterized by a wing membrane lacking macrotrichia. The subfamily, as well as the family as a whole, is characterized by the presence o
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