Revision and new data on the Early and Middle Miocene soricids (Soricomorpha, Mammalia) from Central and South-Eastern F
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Revision and new data on the Early and Middle Miocene soricids (Soricomorpha, Mammalia) from Central and South-Eastern France Marguerite Hugueney • Pierre Mein Olivier Maridet
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Received: 6 November 2011 / Accepted: 15 December 2011 / Published online: 13 January 2012 Ó Akademie der Naturwissenschaften Schweiz (SCNAT) 2012
Abstract New or poorly known Soricids from the Early Miocene of Central France (Limagne d’Allier) and from the Middle Miocene of South-Eastern France (near Lyon) are studied; one new genus (Viretia) and a new species (Carposorex burkarti) are described. The soricid assemblages of the Early and late Early/Middle Miocene localities are totally different from each other and new information about appearance and evolution of the taxa are introduced. The following are lists of studied taxa for the beginning of the Early Miocene: Clapasorex bonisi, Clapasorex aff. sigei, Carposorex burkarti nov. sp., Soricella cf. discrepans, Oligosorex aff. antiquus and Crocidosoricinae indet, and for the late Early/Middle Miocene Dinosorex sansaniensis, Dinosorex pachygnathus, Heterosorex delphinensis, Miosorex desnoyersianus, Miosorex grivensis, Lartetium dehmi, Lartetium ziegleri, Turiasorex pierremeini, Viretia nov. gen. gracilidens, Paenelimnoecus micromorphus, Paenelimnoecus crouzeli and Hemisorex robustus. Keywords Soricidae Heterosoricinae Crocidosoricinae Allosoricinae Paenelimnoecinae Soricinae Early Neogene France Central Massif Region of Lyon New taxa
M. Hugueney (&) P. Mein O. Maridet Laboratoire de ge´ologie de Lyon: terre, plane`tes, environnement, UMR CNRS 5276, universite´ Lyon 1, 43, boulevard du 11-novembre, 69622 Villeurbanne cedex, France e-mail: [email protected] P. Mein e-mail: [email protected] O. Maridet Key Laboratory of Evolutionary Systematics of Vertebrates, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China e-mail: [email protected]
Introduction The Early and Middle Miocene fossil record of Central and South-Eastern France comprises numerous and often very rich localities, either from stratified sediments or fissurefillings, all of which yielded Soricids. The initial studies of these taxa date back several decades and most of them have never been restudied since then. Furthermore, newly discovered specimens have never been described. We survey and update taxa that belong to the subfamilies: Heterosoricinae, Allosoricinae, Paenelimnoecinae, Soricinae and Crocidosoricinae, the last of which is paraphyletic (as suggested by Furio´ et al. 2007; Hugueney and Maridet 2011 and others) but it is convenient to separate the taxa whose phylogenetic positions are uncertain from those whose assignment is certain. Biochronologic background of Limagne Basin localities The Limagne Basin yielded the types of several soricid genera (Crocidosorex Lavocat, 1952, Oligosorex Kretzoi, 1959, Meingensorex Hugueney and Maridet, 2011; see Fig. 1). The basin includes for instance the well-known loca
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