A new Andean species of Protium . Studies in neotropical Burseraceae XXIX
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The New York Botanical Garden, 2900 Southern Blvd., Bronx, NY 10458–5126, USA; e-mail: [email protected] 2 Facultad de Ciencias Forestales, Universidad Nacional Agraria La Molina, Ap. 459, Lima 12, Peru; e-mail: [email protected]
Abstract. Protium oxapampae is described and illustrated. The Burseraceae are well-known for being a diverse and often dominant group in Amazonia, but this species exemplifies the under-appreciated and to a large extent undescribed diversity of the family in the Andes. Keywords: Biodiversity, montane forests, Peru, taxonomy. Resumen. Protium oxapampae es descrito e ilustrado. Las Burseraceae son bien conocidas por ser un grupo diverso y a menudo dominante en la Amazonía, pero esta especie es un ejemplo de que conforman una familia poco considerada con presencia escasamente descrita para los Andes.
Protium Burm.f. is a pantropical genus, but the vast majority of its species are Neotropical, with one apparently relictual species each in Madagascar, Mauritius, New Guinea, Australia, Indonesia, continental SE Asia, and the Philippines (Daly et al., 2011). Now that Crepidospermum Benth. and Tetragastris Gaertn. have been subsumed under Protium, tribe Protieae is monogeneric and at present comprises ca. 145 published and accepted species (Daly & Fine, 2018 and subsequent additions). This number does not take into consideration the 60 as yet undescribed species of Protium already represented in herbaria; unfortunately, 25 of these species are represented by material that is either incomplete (lacking fruits and/or one gender of flowers) or of poor quality. Protium is well-known for displaying notably high diversity and/or abundance in lowland terra firme forests, especially Amazonia (e.g., Daly, 2018), but what is most remarkable about the 60 undescribed species is that 28 of them are montane, ranging from 800 up to 2600 meters elevation. The majority of the specimens are the result of rather intense botanical exploration of the Andes during the 1980s, and to date, most species appear to be rather rare and restricted in their
distributions, but that hypothesis may change after more thorough documentation of the Andean tree flora. One of the undescribed species that is most widespread and represented by a greater number of complete specimens is described herein.
Materials and methods The authors examined herbarium material at USM and NY, the latter consisting primarily of duplicates sent from MO to the first author as gifts for determination. Measurements were made on dry material except for flowers, which were hydrated and softened in aerosol and transferred to 70% ETOH to prevent excess softening and facilitate dissection. Leaf architecture terminology follows Ellis et al. (2009). Protium oxapampae Daly & Reynel, sp. nov. Type: Peru. Pasco: 3 km E of Oxapampa, 10°34’S, 75°24’W, elev. 1900–1950 m, 23 Nov 1982 (♂ fl), David N. Smith & Willibaldo Brack-Egg 2809 (holotype: NY-04247940; isotype: MO). (Fig. 1.)
Brittonia, XX(X), 2020, pp. 1–6 © 2020, by The New York Botanical Garden Press
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