First record of Karydomys (Rodentia, Mammalia) from the German part of the North Alpine Foreland Basin

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First record of Karydomys (Rodentia, Mammalia) from the German part of the North Alpine Foreland Basin Je´roˆme Prieto • Herbert Scholz

Received: 11 March 2013 / Accepted: 12 September 2013 / Published online: 16 November 2013 Ó Swiss Geological Society 2013

Abstract Karydomys, a rare genus of cricetid rodents from the Middle Miocene of central Europe, had previously been reported only from the Swiss part of the North Alpine Foreland basin; documented evidence from the German part was lacking. This paper describes several new specimens of Karydomys from the German localities Ho¨ll and Laimering 3. A correlation of both localities to the Bavarian local biostratigraphic scale OSM F is proposed. Taxonomically, the fossils are most probably linked to K. wigharti from Hambach 6C (north-west Germany), and thus assigned to K. cf. wigharti. In spite of the scarcity of Karydomys fossils in the Upper Freshwatermolasse, the taxon is an important biostratigraphical marker because of its short stratigraphical range.

Keywords Cricetid rodent  Vertebrate palaeontology  Upper Freshwatermolasse  Biostratigraphy  Middle Miocene Abbreviations BSPG Bayerische Staatssammlung fu¨r Pala¨ontologie und Geologie, Munich, Germany NRMZ Naturkundemuseum und Ro¨misches Museum im Zumsteinhaus in Kempten NAFB North Alpine Foreland Basin OSM Upper Freshwater molasse from the German OSM (Obere Su¨ßwassermolasse), used herein for the local biostratigraphic units

1 Introduction

Editorial handling: D. Becker & D. Marty. J. Prieto Institute for Geoscience, Eberhard-Karls University, Sigwartstr. 10, 72076 Tu¨bingen, Germany J. Prieto (&) Department for Earth and Environmental Sciences, Ludwig-Maximilians-University Munich and Bavarian State Collections for Palaeontology and Geology, Richard-Wagner-Strasse 10, 80333 Munich, Germany e-mail: [email protected] H. Scholz UBVGEO Lehrstuhl fu¨r Ingenieurgeologie (Prof. Thuro), Technische Universita¨t Mu¨nchen, Arcisstraße 21, 80333 Munich, Germany

Documented evidence from Central Europe of the cricetid genus Karydomys is exceedingly rare. Although several hundred Miocene vertebrate fossil localities are known from the North Alpine Foreland Basin (NAFB; e.g. Ka¨lin and Kempf 2009; Abdul Aziz et al. 2010), the published record of Karydomys is restricted to a few isolated teeth from Middle Miocene localities in Switzerland (e.g., Garapich and Ka¨lin 1999; Bolliger 2000; Ka¨lin and Kempf 2009). Moreover, the genus has been reported from several fissure fillings in the Franconian Alb, Germany (Garapich and Ka¨lin 1999; Mo¨rs and Kalthoff 2004; Prieto 2012). A large tooth sample from the Lower Rhine Embayment (Northwest Germany) was described as a new species by Mo¨rs and Kalthoff (2004). However, the occurrence of Karydomys in the German part of the NAFB had not been demonstrated to date. In this paper, we describe remains of Karydomys from Ho¨ll and Laimering 3, Bavaria. These

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fossils represent the first documented evidence of the genus from the German part of the NAFB.

2 Geographical an