Population Genetics
POPULATION GENETICS IS OFTEN THOUGHT TO BE A DIFFICULT SUBJECT. To some extent, difficulties are inevitable in a field where some quite basic points are controversial. However, problems are most acute when theoretical points are discussed, despite the fac
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TERTIARY L.EVEL BIOLOGY
A series covering selected areas of biology at advanced undergraduate level. While designed specifically for course options at this level within Universities and Polytechnics, the series will be of great value to specialists and research workers in other fields who require a knowledge of the essentials of a subject.
Titles in the series: Biological Membranes Water and Plants Comparative Immunobiology Methods in Experimental Biology Experimentation in Biology Visceral Muscle An Introduction to Biological Rhythms Biology of Nematodes Biology of Ageing Biology of Reproduction An Introduction to Marine Science Biology of Fresh Waters An Introduction to Developmental Biology Neurosecretion Physiology of Parasites Biology of Communication
Harrison and Lunt Meidner and Sheriff Manning and Turner Ralph Ridgman Huddart and Hunt Saunders Croll and Matthews Lamb Hogarth Meadows and Campbell Maitland Ede Maddrell and Nordmann Chappell Lewis and Gower
TERTIARY LEVEL BIOLOGY
Population Genetics
J. S. GALE Lecturer in Genetics University of Birmingham England
Blackie Glasgow and London
Blackie & Son Limited Bishopbriggs Glasgow G64 2NZ Furnival House 14-18 High Holborn London We1 V 6BX
© 1980 J. S. Gale Softcover reprint of the hardcover 1st edition 1980 First published 1980
lSBN-13: 978-1-4613-3926-7 DOl: 10.1007/978-1-4613-3924-3
e-1SBN-13: 978-1-4613-3924-3
All rights reserved. No part of this publication may be reproduced, stored in a retrieval system or transmitted, in any form or by any means, electronic, mechanical, recording or otherwise, without prior permission of the Publishers
International Standard Book Numbers Hardback lSBN-13: 978-1-4613-3926-7 Paperback lSBN-13: 978-1-4613-3924-3
Filmset by Advanced Filmsetters (Glasgow) Limited
Preface
POPULATION GENETICS IS OFTEN THOUGHT TO BE A DIFFICULT SUBJECT.
To some extent, difficulties are inevitable in a field where some quite basic points are controversial. However, problems are most acute when theoretical points are discussed, despite the fact that there has been very little controversy over the mathematics. In my experience, the actual mathematical manipulations rarely cause much difficulty. Rather it is that the biologist, lacking the physicist's or chemist's experience in "reading" mathematical formulae, finds it difficult to appreciate what is happening in a mathematical treatment and to grasp the implications of the results obtained, when these are given in mathematical form. Accordingly, I have followed a procedure, which students seem to find helpful, of giving a rough-and-ready verbal treatment of a problem before attempting a much more exact mathematical treatment; when the results of the latter are not readily interpretable, I have given an elucidation. Another problem which often concerns students is the reliability of results obtained using approximate methods; I have, therefore, discussed this in fair detail in critical cases. When dealing with controversial issues, I have done my very best to be fair. To concea
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