Tropical Montane Cloud Forests

Until relatively recently the valuable tropical montane cloud forests (hereaf­ ter usually referred to as TMCFs) of the world had scarcely come under the assaults experienced by the downslope montane and lowland forests. TMCFs are not hospitable environme

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Edited by O.L. Lange, Wiirzburg, FRG H.A. Mooney, Stanford, USA H. Remmert, Marburg, FRGt

Ecological Studies Volumes published since 1989 are listed at the back of this book.

Lawrence S. Hamilton James O. Juvik F. N. Scatena Editors

Tropical~ontane

Cloud Forests With 102 illustrations

Springer-Verlag New York Berlin Heidelberg London Paris Tokyo Hong Kong Barcelona Budapest

F. N. Scatena International Institute of Tropical Forestry USDA Forest Service Rio Piedras, PR 00928 USA

Lawrence Hamilton Islands and Highlands Environmental Consultancy RR #1, Box l685A Hinesburg, VT 05461 USA James o. Juvik Department of Geography University of Hawaii, Hilo Hilo, HI 96720 USA

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EAST-WEST CENTER PROGRAM ON ENVIRONMENT

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International Hydrological Programme

International Institute of Tropical Forestry

Library of Congress Cataloging-in-Publication Data Tropical montane cloud forests / Lawrence S. Hamilton, James o. Juvik, and F. N. Scatena, editors. p. cm. - (Ecological studies; v. 110) Includes bibliographical references and index. ISBN-13:978-1-4612-7564-0 l. Cloud forest ecology. 2. Cloud forests. 3. Mountain ecologyTropics. 1. Hamilton, Lawrence S. II. Juvik, James O. m. Scatena, F. N. IV. Series. QH54l.5.C63T76 1994 94-12683 574.5'2642-ilc20 Printed on acid-free paper. An earlier version of this book was published by the East-West Center.

@1995 Springer-Verlag New York, Inc. Softcover reprint of the hardcover 1st edition 1995 All rights reserved. This work may not be translated or copied in whole or in part without the written permission of the publisher (Springer-Verlag New York, Inc., 175 Fifth Avenue, New York, NY 10010, USA), except for brief excerpts in connection with reviews or scholarly analysis. Use in connection with any form of information storage and retrieval, electronic adaptation, computer software, or by similar or dissimilar methodology now known or hereafter developed is forbidden. The use of general descriptive names, trade names, trademarks, etc., in this publication, even if the former are not especially identified, is not to be taken as a sign that such names, as understood by the Trade Marks and Merchandise Marks Act, may accordingly be used freely by anyone. Permission to photocopy for internal or personal use, or the internal or personal use of specific clients, is granted by Springer-Verlag, Inc., for libraries registered with the Copyright Clearance Center (CCC), provided that the base fee of $5.00 per copy, plus $0.20 per page, is paid directly to Springer-Verlag New York, 175 Fifth Avenue, New York, NY 10010, USA. ISBN-13:978-1-4612-7564-0 1995 $5.00 + 0.20 Production managed by Laura Carlson; manufacturing supervised by Jacqui Ashri. Compostion by KP Company, Brooklyn, NY. 9 876 543 2 I ISBN-13:978-1-4612-7564-0 e- ISBN-13:978-1-4612-2500- 3 DOl: 10.1007/978-1-4612-2500-3

In Memory of Alwyn Gentry The editors dedicate these proceedings to Dr. Alwyn Gentry, a participant in the symposium and a stalwart champion for tropical cloud forest conservation. His untimely