Wild Emmer, Triticum dicoccoides,Wheat Progenitor: Origin and Evolution

Genetic and morphological evidence clearly indicates that the cultivated tetraploid turgidum wheats (both hulled dicoccum forms and free threshing durum varieties) are closely related to the wild wheat that is native to the Near East and traditionally cal

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Springer-Verlag Berlin Heidelberg GmbH

E. Nevo . A.B. Korol . A. Beiles . T. Fahima

Evolution of Wild Emmer and Wheat Improvement Population Genetics, Genetic Resources, and Genome Organization of Wheat's Progenitor, Triticum dicoccoides

With 59 Figures and 85 Tables

Springer

Professor EVIATAR NEVO Professor ABRAHAM B. KOROL Dr. AVIGDOR BElLES Dr. TZION FAHIMA Institute of Evolution University of Haifa Mount Carmel Haifa 31905 Israel

ISBN 978-3-642-07512-4 Library of Congress Cataloging-in-Publication Data Nevo, Eviatar. Evolution of Wild emmer and wheat improvement: population genetics, genetic resources, and genome organization of wheat's projenitor, Triticum dicoccoides I E. Nevo, A.B. Korol, T. Fahima. p. cm. Includes bibliographical references (p. ). ISBN 978-3-642-07512-4 ISBN 978-3-662-07140-3 (eBook) DOI 10.1007/978-3-662-07140-3 1. Emmer wheat - Evolution.

Bentsionovich)

2. Crop improvement. I. Korol', A.B. (Abram II. Fahima, T. (Tzion), 1959- III. Title.

This work is subject to copyright. All rights reserved, whether the whole or part of the material is concerned, specifically the rights of translation, reprinting, reuse of illustrations, recitation, broadcasting, reproduction on microfilm or in any other way, and storage in data banks. Duplication of this publication or parts thereof is permitted only under the provisions of the German Copyright Law of September 9, 1965, in its current version, and permission for use must always be obtained from Springer-Verlag Berlin Heidelberg GmbH. Violations are liable for prosecution under the German Copyright Law. http://www.springer.de © Springer-Verlag Berlin Heidelberg 2002 Originally published by Springer-Verlag Berlin Heidelberg New York in 2002 Sof'tcover reprint of the hardcover 1st edition 2002

The use of general descriptive names, registered names, trademarks, etc. in this publication does not imply, even in the absence of a specific statement, that such names are exempt from the relevant protective laws and regulations and therefore free for general use. Cover design: D & P, Heidelberg Cover photo: Wild Emmer wheat, Triticum dicoccoides, Almagor, North of the Sea of Galilee, Israel (Photo by Tzion Fahima) Typesetting: Best-set Typesetter Ltd., Hong Kong SPIN 10795445 31/3130-5432 1 0 - Printed on acid-free paper

Preface

This book is about the contribution to evolutionary theory and agricultural technology of one of humankind's most dramatic imitations of the evolutionary process, namely crop domestication, as exemplified by the progenitor of wheat, Triticum dicoccoides. This species is a major model organism and it has been studied at the Institute of Evolution, University of Haifa, since 1979. The domestication by humans of wild plants to cultivated ones during the last ten millennia is one of the best demonstrations of evolution. It is a process that has been condensed in time and advanced by artificial rather than natural selection. Plant and animal domestication revolutionized human cultural evolution and is the major factor underlying