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Biotechnology in Agriculture and Forestry Volumes already published and in preparation are listed at the end of this book.

Biotechnology in Agriculture and Forestry 57 Plant Metabolomics Edited by K. Saito, R.A. Dixon, and L. Willmitzer

With 96 Figures, 29 in Color, and 10 Tables

123

Series Editors Professor Dr. Toshiyuki Nagata University of Tokyo Graduate School of Science Department of Biological Sciences 7-3-1 Hongo, Bunkyo-ku Tokyo 113-0033, Japan Professor Dr. Horst Lörz Universität Hamburg Biozentrum Klein Flottbek Zentrum für Angewandte Molekularbiologie der Pflanzen (AMP II) Ohnhorststraße 18 22609 Hamburg, Germany

Professor Dr. Jack M. Widholm University of Illinois 285A E.R. Madigan Laboratory Department of Crop Sciences 1201 W. Gregory Urbana, IL 61801, USA

Volume Editors Professor Dr. Kazuki Saito Chiba University Graduate School of Pharmaceutical Sciences Yayoi-cho 1-33, Inage-ku Chiba 263-8522, Japan; RIKEN Plant Science Center Yokohama 230-0045, Japan Professor Dr. Richard A. Dixon Plant Biology Division Samuel Roberts Noble Foundation 2510 Sam Noble Parkway Ardmore, OK 73401, USA

Professor Dr. Lothar Willmitzer Max Planck Institute of Molecular Plant Physiology Am Mühlenberg 1 14476 Golm, Germany

Library of Congress Control Number: 2005936763

ISSN 0934-943X ISBN-10 3-540-29781-2 Springer Berlin Heidelberg New York ISBN-13 978-3-540-29781-9 Springer Berlin Heidelberg New York This work is subject to copyright. All rights reserved, whether the whole or part of the material is concerned, specifically the rights of translation, reprinting, reuse of illustrations, recitation, broadcasting, reproduction on microfilm or in any other way, and storage in data banks. Duplication of this publication or parts thereof is permitted only under the provisions of the German Copyright Law of September 9, 1965, in its current version, and permission for use must always be obtained from Springer. Violations are liable for prosecution under the German Copyright Law. Springer is a part of Springer Science + Business Media springer.com © Springer-Verlag Berlin Heidelberg 2006 Printed in Germany The use of general descriptive names, registered names, trademarks, etc. in this publication does not imply, even in the absence of a specific statement, that such names are exempt from the relevant protective laws and regulations and therefore free for general use. Editor: Dr. Dieter Czeschlik, Heidelberg, Germany Desk Editor: Dr. Andrea Schlitzberger, Heidelberg, Germany Cover design: design&production GmbH, Heidelberg, Germany Typesetting and production: LE-TEX Jelonek, Schmidt & Vöckler GbR, Leipzig, Germany Printed on acid-free paper 31/3152 5 4 3 2 1 0

Preface

Metabolomics is a rapidly-emerging sector of post-genome research. The metabolome (a set of all metabolites of an organism) represents not only the ultimate phenotype of cells by the perturbation of gene expression and the modulation of protein functions caused by the environment or mutations, but the metabolome can also feed back on gene expression and prote