New and Little Known Gryllones Insects (Insecta: Gryllones: Cnemidolestida, Reculida) from the Triassic of Eurasia
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and Little Known Gryllones Insects (Insecta: Gryllones: Cnemidolestida, Reculida) from the Triassic of Eurasia D. S. Aristova, b, * a
Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow, Russia bCherepovets State University, Cherepovets, Russia *e-mail: [email protected] Received November 7, 2019; revised November 18, 2019; accepted November 18, 2019
Abstract—New and little known Gryllones insects (Insecta: Gryllones) from the Triassic of Eurasia are described. The genus Yontala Aristov, 2005 from the Lower Triassic locality of Yontala (Induan Stage of the Vologda Region, Russia) is re-described and transferred to the family Sylvabestiidae of the order Cnemidolestida. Sharovala triassica gen. et sp. nov. (Cnemidolestida: Sylvabestiidae) from the Middle Triassic Madygen locality (Ladinian Stage of the Osh region of Kyrgyzstan) is described. New representatives of the order Reculida, Chauliodites sakmaris sp. nov. (family Chaulioditidae) from the Lower Triassic locality of Petropavlovka (Olenekian Stage of the Orenburg Region, Russia) and Batkenopterum kirgizicum gen. et sp. nov. (family Sylvaphlebiidae) from Madygen are described. Changes in Cnemidolestida and Reculida faunas at the boundary of the Paleozoic and Mesozoic are discussed. Keywords: insects, Cnemidolestida, Reculida, Triassic, Eurasia DOI: 10.1134/S0031030120050020
INTRODUCTION Changes in the faunas of some orders of Gryllones insects (Cnemidolestida and Reculida) in the time interval close to the boundary of the Permian and Triassic are discussed in this paper. The interval that extends from the Vyatkian Stage of the Upper Permian to the Ladinian Stage of the Triassic (Madygen locality) is considered. The distribution of the families of the orders discussed is inferred from the studies (Storozhenko, 1998; Rasnitsyn et al., 2013; Aristov, 2014, 2018a, b) for the Permian and from the studies (Storozhenko, 1998; Aristov, 2003, 2020; Aristov and et al., 2011; Bashkuev et al., 2011; Aristov, 2014, 2018a, b) for the Triassic, as well as from the author’s original data. Cnemidolestida and Reculida faunas from a range of localities (Fig. 1) are considered in this study: the Upper Permian localities include the Vyatkian Belmont (Australia, New South Wales), Aristovo (Russia, Vologda Region), Sokovka (Russia, Vladimir Region), Nedubrovo (Russia, Vologda Region), Babii Kamen (Russia, Kemerovo Region), Tunguska (a group of localities in the Krasnoyarsk Region of Russia), Zalazna (Russia, Kirov Region) and YamanUs (Mongolia, South Gobi Aymag: Uranbileg, 2019). Yontala (Russia, Vologda Region) represents the Induan Stage of the Lower Triassic. The localities of Tikhvinskoe (Russia, Yaroslavl Region) and Petropavlovka (Russia, Orenburg Region) date back to the
Olenekian age, whereas Hammelburg and others (a group of localities in Germany, Lower Franconia, and Thuringia) and Vosges (a group of localities in France, Alsace, and Lorraine) represent the Anisian Stage, and Madygen (Kyrgyzstan, Osh region) is assigned to the Ladinian layer of the
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